Mihai's Paleobotany Chronicles

Pachypteris gradinarui POPA 2000
2000 Pachypteris gradinarui POPA, p. 33-46, Text-figs. 1-5, Pls. 1-3.
2004 Pachypteris gradinarui GUIGNARD, POPA and BARALE, p. 263-273, Pl. 1.

Pteridospermopsida/Corystospermales/Incertae sedis

Bipinnate leaves, rachis longitudinally striated, with a long petiole and secondary rachises inserted oppositely to suboppositely on the upper side of the primary rachis. Pinnules rounded to elongated, with contracted anadromic and decurrent catadromic margins, always inserted on the upper side of rachis, often overlapping, without any marginal, abaxial thickening or folding, margins entire, apex atenuated, apical pinnules in a pinna enlarged. Intercalary pinnules occur occasionally. Pinnule midrib prominent abaxially; secondary veins macroscopically invisible. Leaves hypostomatic, the upper cuticle thicker than the lower one. Upper cuticle with polygonal, rounded epidermal cells, no indication of secondary veins, no papillae. Lower cuticle with a well-defined midrib represented by rows of slightly elongated cells, secondary veins indicated near the midrib but not further to the margin, sometimes showing a single dichotomy. Stomata haplocheilic, monocyclic or incompletely dicyclic, scattered all over the lower epidermal surface except the midrib, with sunken and elongated guard cells, 4-7 trapezoid subsidiary cells. In Popa (2000).

P. gradinarui has an intresting position within the Pachypteris group, somehow intermediary between Pachypteris and Cycadopteris, but it lacks clear marginal thickenings or folds that characterize the latter genus.

Cristian, Getic Nappe, South Carpathians.

Sinemurian in age.


The degree of preservation is very good.

Popa, M.E., 2000. Aspects of Romanian Early Jurassic palaeobotany and palynology. Part II. A new species of Pachypteris from Cristian. Review of Palaeobotany and Palynology, 111: 31-47.

Guignard, G., Popa, M.E. and Barale, G., 2004. Ultrastructure of Early Jurassic fossil plant cuticles: Pachypteris gradinarui POPA. Tissue and Cell: 263-273.

Pachypteris gradinarui, holotype, sample G33, fragment 1, Cristian.